CAMPBELL
BIOLOGY TWELFTH EDITION
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Lisa A. Urry Michael L. Cain Steven A. Wasserman
MILLS COLLEGE, OAKLAND, NEW MEXICO STATE UNIVERSITY UNIVERSITY OF CALIFORNIA,
CALIFORNIA SAN DIEGO
Peter V. Minorsky Rebecca B. Orr
MERCY COLLEGE, DOBBS FERRY, COLLIN COLLEGE, PLANO, TEXAS
NEW YORK
URRY8743_12_SE_FM_PRF.indd 1
, Director, Global Higher Ed Content Management and Project Manager: Katie Cook
Strategy, Science & Health Sciences: Jeanne Zalesky Content Producers, Mastering Biology: Kaitlin Smith,
Manager, Higher Ed Global Content Strategy, Life Ashley Gordon
Sciences: Joshua Frost Supervising Media Producer: Tod Regan
Associate Content Analyst: Chelsea Noack Media Producer: Ziki Dekel
Editorial Assistant: Ashley Fallon Full-Service Vendor: Integra Software Services, Inc.
Director, Higher Ed Product Management, Design Manager: Mark Ong
Life Sciences: Michael Gillespie Cover & Interior Designer: Jeff Puda
Product Manager: Rebecca Berardy Schwartz Illustrators: Lachina Creative
Managing Producer: Michael Early Rights & Permissions Project Manager: Matt Perry,
Senior Content Producer: Lori Newman SPi Global
Director, Content Development & Partner Rights & Permissions Manager: Ben Ferrini
Relationships: Ginnie Simione Jutson Photo Researcher: Maureen Spuhler
Supervising Editors: Beth N. Winickoff, Pat Burner Product and Solutions Specialist: Kelly Galli
Senior Developmental Editors: John Burner, Mary Ann Senior Product Marketing Manager: Alysun Estes
Murray, Hilair Chism, Andrew Recher, Mary Hill Manufacturing Buyer: Stacey Weinberger, LSC
Specialist, Instructional Design and Development: Communications
Sarah Young-Dualan Cover Photo Credit: Robert Rohrbaugh/Rohrbaugh
Senior Content Developer, Mastering Biology: Photography
Sarah Jensen
Copyright © 2021, 2017, 2014 by Pearson Education, Inc. or its affiliates, 221 River Street, Hoboken, NJ 07030. All
a
Rights Reserved. Manufactured in the United States of America. This publication is protected by copyright, and
y
permission should be obtained from the publisher prior to any prohibited reproduction, storage in a retrieval
system, or transmission in any form or by any means, electronic, mechanical, photocopying, recording, or
i
otherwise. For information regarding permissions, request forms, and the appropriate contacts within the
gh
Pearson Education Global Rights and Permissions department, please visit www.pearsoned.com/permissions/.
Acknowledgments of third-party content appear in the Credits section (beginning on page CR-1), which
constitutes an extension of this copyright page.
gt
PEARSON, ALWAYS LEARNING, MasteringTM Biology, and BioFlix® are exclusive trademarks owned by Pearson
u
Education, Inc. or its affiliates in the U.S. and/or other countries.
a
Unless otherwise indicated herein, any third-party trademarks that may appear in this work are the property of their
b
respective owners and any references to third-party trademarks, logos or other trade dress are for demonstrative or
descriptive purposes only. Such references are not intended to imply any sponsorship, endorsement, authorization,
M
or promotion of Pearson’s products by the owners of such marks, or any relationship between the owner and
Pearson Education, Inc. or its affiliates, authors, licensees or distributors.
M
Library of Congress Cataloging-in-Publication Data
Names: Urry, Lisa A., author. | Cain, Michael L. (Michael Lee), author. |
Wasserman, Steven Alexander, author. | Minorsky, Peter V., author. |
Orr, Rebecca B., author. | Campbell, Neil A., Biology.
Title: Campbell biology / Lisa A. Urry, Michael L. Cain, Steven A.
Wasserman, Peter V. Minorsky, Rebecca B. Orr, Neil A. Campbell.
Description: Twelfth edition. | New York, NY : Pearson, 2020. | Includes index.
Identifiers: LCCN 2019039139 | ISBN 9780135188743 (hardcover) |
ISBN 9780135988046 (ebook)
Subjects: LCSH: Biology.
Classification: LCC QH308.2 .C34 2020 | DDC 570—dc23
LC record available at https://lccn.loc.gov/2019039139
ScoutAutomatedPrintCode
ISBN 10: 0-135-18874-1; ISBN 13: 978-0-135-18874-3 (Rental Edition)
ISBN 10: 0-136-62344-1; ISBN 13: 978-0-136-62344-1 (Instructor Review Copy)
www.pearson.com
URRY8743_12_SE_FM_PRF.indd 2
, Setting the Standard for Excellence,
Accuracy, and Innovation
Campbell Biology, 12th Edition, delivers an authoritative, accurate, current,
and pedagogically innovative experience that helps students make connections
so they learn and understand biology. This edition presents new, engaging visual
and digital resources that meet demonstrated student needs.
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, A New Visual Experience for
Every Chapter
NEW! Chapter Openers introduce each chapter and feature a question
answered with a clear, simple image to help students visualize and remember
concepts as they move through each chapter. Each opener includes a Study
Tip and highlights of interactive media in Mastering Biology.
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NEW! A Visual
Overview helps
students start with the
big picture.
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URRY8743_12_SE_FM_PRF.indd 4
BIOLOGY TWELFTH EDITION
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Lisa A. Urry Michael L. Cain Steven A. Wasserman
MILLS COLLEGE, OAKLAND, NEW MEXICO STATE UNIVERSITY UNIVERSITY OF CALIFORNIA,
CALIFORNIA SAN DIEGO
Peter V. Minorsky Rebecca B. Orr
MERCY COLLEGE, DOBBS FERRY, COLLIN COLLEGE, PLANO, TEXAS
NEW YORK
URRY8743_12_SE_FM_PRF.indd 1
, Director, Global Higher Ed Content Management and Project Manager: Katie Cook
Strategy, Science & Health Sciences: Jeanne Zalesky Content Producers, Mastering Biology: Kaitlin Smith,
Manager, Higher Ed Global Content Strategy, Life Ashley Gordon
Sciences: Joshua Frost Supervising Media Producer: Tod Regan
Associate Content Analyst: Chelsea Noack Media Producer: Ziki Dekel
Editorial Assistant: Ashley Fallon Full-Service Vendor: Integra Software Services, Inc.
Director, Higher Ed Product Management, Design Manager: Mark Ong
Life Sciences: Michael Gillespie Cover & Interior Designer: Jeff Puda
Product Manager: Rebecca Berardy Schwartz Illustrators: Lachina Creative
Managing Producer: Michael Early Rights & Permissions Project Manager: Matt Perry,
Senior Content Producer: Lori Newman SPi Global
Director, Content Development & Partner Rights & Permissions Manager: Ben Ferrini
Relationships: Ginnie Simione Jutson Photo Researcher: Maureen Spuhler
Supervising Editors: Beth N. Winickoff, Pat Burner Product and Solutions Specialist: Kelly Galli
Senior Developmental Editors: John Burner, Mary Ann Senior Product Marketing Manager: Alysun Estes
Murray, Hilair Chism, Andrew Recher, Mary Hill Manufacturing Buyer: Stacey Weinberger, LSC
Specialist, Instructional Design and Development: Communications
Sarah Young-Dualan Cover Photo Credit: Robert Rohrbaugh/Rohrbaugh
Senior Content Developer, Mastering Biology: Photography
Sarah Jensen
Copyright © 2021, 2017, 2014 by Pearson Education, Inc. or its affiliates, 221 River Street, Hoboken, NJ 07030. All
a
Rights Reserved. Manufactured in the United States of America. This publication is protected by copyright, and
y
permission should be obtained from the publisher prior to any prohibited reproduction, storage in a retrieval
system, or transmission in any form or by any means, electronic, mechanical, photocopying, recording, or
i
otherwise. For information regarding permissions, request forms, and the appropriate contacts within the
gh
Pearson Education Global Rights and Permissions department, please visit www.pearsoned.com/permissions/.
Acknowledgments of third-party content appear in the Credits section (beginning on page CR-1), which
constitutes an extension of this copyright page.
gt
PEARSON, ALWAYS LEARNING, MasteringTM Biology, and BioFlix® are exclusive trademarks owned by Pearson
u
Education, Inc. or its affiliates in the U.S. and/or other countries.
a
Unless otherwise indicated herein, any third-party trademarks that may appear in this work are the property of their
b
respective owners and any references to third-party trademarks, logos or other trade dress are for demonstrative or
descriptive purposes only. Such references are not intended to imply any sponsorship, endorsement, authorization,
M
or promotion of Pearson’s products by the owners of such marks, or any relationship between the owner and
Pearson Education, Inc. or its affiliates, authors, licensees or distributors.
M
Library of Congress Cataloging-in-Publication Data
Names: Urry, Lisa A., author. | Cain, Michael L. (Michael Lee), author. |
Wasserman, Steven Alexander, author. | Minorsky, Peter V., author. |
Orr, Rebecca B., author. | Campbell, Neil A., Biology.
Title: Campbell biology / Lisa A. Urry, Michael L. Cain, Steven A.
Wasserman, Peter V. Minorsky, Rebecca B. Orr, Neil A. Campbell.
Description: Twelfth edition. | New York, NY : Pearson, 2020. | Includes index.
Identifiers: LCCN 2019039139 | ISBN 9780135188743 (hardcover) |
ISBN 9780135988046 (ebook)
Subjects: LCSH: Biology.
Classification: LCC QH308.2 .C34 2020 | DDC 570—dc23
LC record available at https://lccn.loc.gov/2019039139
ScoutAutomatedPrintCode
ISBN 10: 0-135-18874-1; ISBN 13: 978-0-135-18874-3 (Rental Edition)
ISBN 10: 0-136-62344-1; ISBN 13: 978-0-136-62344-1 (Instructor Review Copy)
www.pearson.com
URRY8743_12_SE_FM_PRF.indd 2
, Setting the Standard for Excellence,
Accuracy, and Innovation
Campbell Biology, 12th Edition, delivers an authoritative, accurate, current,
and pedagogically innovative experience that helps students make connections
so they learn and understand biology. This edition presents new, engaging visual
and digital resources that meet demonstrated student needs.
yi a
gtgh
bau
M
M
URRY8743_12_SE_FM_PRF.indd 3
, A New Visual Experience for
Every Chapter
NEW! Chapter Openers introduce each chapter and feature a question
answered with a clear, simple image to help students visualize and remember
concepts as they move through each chapter. Each opener includes a Study
Tip and highlights of interactive media in Mastering Biology.
yi a
gtgh
bau
M
M
NEW! A Visual
Overview helps
students start with the
big picture.
iv
URRY8743_12_SE_FM_PRF.indd 4
