STUDENT C O M P A N I O N
F U N D A M
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E N T A L S O F
Biochemistry
LIFEAT n
THE MOLECULAR
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LEVEL n
Voet Voet Pratt
F O U R T H E D I T I O N
Akif Uzman Jerry Johnson Joseph Eichberg William Widger
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Donald Voet Judith G. Voet Charlotte W. Pratt
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FUNDAMENTALS OF n
BIOCHEMISTRY
LIFE AT THE MOLECULAR LEVEL
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Fourth Edition n
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University of Houston n n
Jerry Johnson
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University of Houston n n
Joseph Eichberg n
University of Houston n n
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Swarthmore College n
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JOHN WILEY & SONS, INC
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Cover Illustration Norm Christiansen
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Cover Photos
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(Vitruvian Man): © Odysseus/Alamy and © Dennis Hallinan/Alamy n n n n n n n
Molecular structures clockwise from top: based on X-ray structures that were respectively determined by
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Richard Dickerson and Horace Drew, Caltech; Gerard Bunick, University of Tennessee; Thomas Steitz,
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Yale University; Alfonso Mondragón, Northwestern University; Venki Ramakrishnan, MRC Laboratory
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of Molecular Biology, Cambridge, U.K.; Andrew Leslie and John Walker, MRC Laboratoryof Molecular
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Biology, Cambridge, U.K.
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Student Companion (paperback) n n ISBN-13 978-1-1182-1827-3 n
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n n n n n n n n n n
F U N D A M
n
E N T A L S O F
Biochemistry
LIFEAT n
THE MOLECULAR
n
LEVEL n
Voet Voet Pratt
F O U R T H E D I T I O N
Akif Uzman Jerry Johnson Joseph Eichberg William Widger
n n n n
Donald Voet Judith G. Voet Charlotte W. Pratt
n n n n n n
,This page is intentionally left blank
n n n n n
,STUDENT COMPANION TO ACCOMPANY n n n
FUNDAMENTALS OF n
BIOCHEMISTRY
LIFE AT THE MOLECULAR LEVEL
n n n n
Fourth Edition n
Akif Uzman
n
University of Houston n n
Jerry Johnson
n
University of Houston n n
Joseph Eichberg n
University of Houston n n
William Widger n
University of Houston n n
Donald Voet n
University of Pennsylvania n n
Judith G. Voet n n
Swarthmore College n
Charlotte W. Pratt n n
Seattle, Washington n
JOHN WILEY & SONS, INC
n n n n
, Cover Designer Madelyn Lesure n n n
Cover Illustration Norm Christiansen
n n n n
Cover Photos
n n
(Vitruvian Man): © Odysseus/Alamy and © Dennis Hallinan/Alamy n n n n n n n
Molecular structures clockwise from top: based on X-ray structures that were respectively determined by
n n n n n n n n n n n n n
Richard Dickerson and Horace Drew, Caltech; Gerard Bunick, University of Tennessee; Thomas Steitz,
n n n n n n n n n n n n n
Yale University; Alfonso Mondragón, Northwestern University; Venki Ramakrishnan, MRC Laboratory
n n n n n n n n n n
of Molecular Biology, Cambridge, U.K.; Andrew Leslie and John Walker, MRC Laboratoryof Molecular
n n n n n n n n n n n n n n
Biology, Cambridge, U.K.
n n n
Founded in 1807, John Wiley & Sons, Inc. has been a valued source of knowledge and understanding for more
n n n n n n n n n n n n n n n n n n
than 200 years, helping people around the world meet their needs and fulfill their aspirations. Our company is
n n n n n n n n n n n n n n n n n n
built on a foundation of principles that include responsibility to the communities we serve andwhere we live
n n n n n n n n n n n n n n n n n n
and work. In 2008, we launched a Corporate Citizenship Initiative, a global effort to address the
n n n n n n n n n n n n n n n n
environmental, social, economic, and ethical challenges we face in our business. Among the issues we are
n n n n n n n n n n n n n n n n
addressing are carbon impact, paper specifications and procurement, ethical conduct within our business and
n n n n n n n n n n n n n n
among our vendors, and community and charitable support. For more information, please visit our website:
n n n n n n n n n n n n n n n
www.wiley.com/go/citizenship.
n
Copyright © 2012 n n John Wiley & Sons, Inc.
n n n n
All rights reserved. No part of this publication may be reproduced, stored in a retrieval system or transmitted
n n n n n n n n n n n n n n n n n
in any form or by any means, electronic, mechanical, photocopying, recording, scanning or otherwise,
n n n n n n n n n n n n n n
except as permitted under Sections 107 or 108 of the 1976 United States Copyright Act, without either the
n n n n n n n n n n n n n n n n n n
prior written permission of the Publisher, or authorization through payment of the appropriate per-copy fee
n n n n n n n n n n n n n n n
to the Copyright Clearance Center, Inc., 222 Rosewood Drive, Danvers, MA 01923, website
n n n n n n n n n n n n n
www.copyright.com. Requests to the Publisher for permission should be addressed to thePermissions
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Department, John Wiley & Sons, Inc. 111 River Street, Hoboken, NJ 07030-5774, (201) 748-6011, fax
n n n n n n n n n n n n n n n
(201) 748-6008, website http://www.wiley.com/go/permissions.
n n n n
To order books or for customer service, please call 1-800-CALL WILEY (225-5945).
n n n n n n n n n n n
Student Companion (paperback) n n ISBN-13 978-1-1182-1827-3 n
Printed in the United States of America n n n n n n
10 9 8 7 6 5 4 3 2 1
n n n n n n n n n n
