,
, Principles of
ANATOMY & PHYSIOLOGY
15th Edition
GERARD J. TORTORA
Bergen Community College
BRYAN DERRICKSON
Valencia College
, VICE PRESIDENT AND DIRECTOR Petra Recter
SENIOR EDITOR Maria Guarascio
ASSOCIATE DEVELOPMENT EDITOR Laura Rama
ASSISTANT DEVELOPMENT EDITOR Lindsey Myers
EDITORIAL ASSISTANT MaryAlice Skidmore
SENIOR MARKETING MANAGER Alan Halfen
SENIOR CONTENT MANAGER Svetlana Barskaya
SENIOR PRODUCTION EDITOR Trish McFadden
SENIOR PHOTO EDITOR MaryAnn Price
SENIOR PRODUCT DESIGNER Linda Muriello
TEXT AND COVER DESIGNER Thomas Nery
COVER PHOTO ©PhotoAlto sas/Alamy Stock Photo
This book was set in 9.5/12.5 Source Sans Pro by Aptara®, Inc. Printed and bound by Quad Graphics. This book is printed on acid-free
paper. ∞
Founded in 1807, John Wiley & Sons, Inc. has been a valued source of knowledge and understanding for more than 200 years, helping
people around the world meet their needs and fulfill their aspirations. Our company is built on a foundation of principles that include
responsibility to the communities we serve and where we live and work. In 2008, we launched a Corporate Citizenship Initiative, a
global effort to address the environmental, social, economic, and ethical challenges we face in our business. Among the issues we are
addressing are carbon impact, paper specifications and procurement, ethical conduct within our business and among our vendors,
and community and charitable support. For more information, please visit our website: www.wiley.com/go/citizenship.
Copyright © 2017 John Wiley & Sons, Inc. All rights reserved.
No part of this publication may be reproduced, stored in a retrieval system, or transmitted in any form or by any means, electronic,
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the Publisher for permission should be addressed to the Permissions Department, John Wiley & Sons, Inc., 111 River Street, Hoboken,
NJ 07030-5774, (201) 748-6011, fax (201) 748-6008, or online at: www.wiley.com/go/permissions.
Evaluation copies are provided to qualified academics and professionals for review purposes only, for use in their courses during the
next academic year. These copies are licensed and may not be sold or transferred to a third party. Upon completion of the review
period, please return the evaluation copy to Wiley. Return instructions and a free-of-charge return shipping label are available at:
www.wiley.com/go/returnlabel. If you have chosen to adopt this textbook for use in your course, please accept this book as your com-
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ePUB ISBN 978-1-119-32064-7
The inside back cover will contain printing identification and country of origin if omitted from this page. In addition, if the ISBN on the
back cover differs from the ISBN on this page, the one on the back cover is correct.
Printed in the United States of America.
10 9 8 7 6 5 4 3 2 1
, Principles of
ANATOMY & PHYSIOLOGY
15th Edition
GERARD J. TORTORA
Bergen Community College
BRYAN DERRICKSON
Valencia College
, VICE PRESIDENT AND DIRECTOR Petra Recter
SENIOR EDITOR Maria Guarascio
ASSOCIATE DEVELOPMENT EDITOR Laura Rama
ASSISTANT DEVELOPMENT EDITOR Lindsey Myers
EDITORIAL ASSISTANT MaryAlice Skidmore
SENIOR MARKETING MANAGER Alan Halfen
SENIOR CONTENT MANAGER Svetlana Barskaya
SENIOR PRODUCTION EDITOR Trish McFadden
SENIOR PHOTO EDITOR MaryAnn Price
SENIOR PRODUCT DESIGNER Linda Muriello
TEXT AND COVER DESIGNER Thomas Nery
COVER PHOTO ©PhotoAlto sas/Alamy Stock Photo
This book was set in 9.5/12.5 Source Sans Pro by Aptara®, Inc. Printed and bound by Quad Graphics. This book is printed on acid-free
paper. ∞
Founded in 1807, John Wiley & Sons, Inc. has been a valued source of knowledge and understanding for more than 200 years, helping
people around the world meet their needs and fulfill their aspirations. Our company is built on a foundation of principles that include
responsibility to the communities we serve and where we live and work. In 2008, we launched a Corporate Citizenship Initiative, a
global effort to address the environmental, social, economic, and ethical challenges we face in our business. Among the issues we are
addressing are carbon impact, paper specifications and procurement, ethical conduct within our business and among our vendors,
and community and charitable support. For more information, please visit our website: www.wiley.com/go/citizenship.
Copyright © 2017 John Wiley & Sons, Inc. All rights reserved.
No part of this publication may be reproduced, stored in a retrieval system, or transmitted in any form or by any means, electronic,
mechanical, photocopying, recording, scanning or otherwise, except as permitted under Sections 107 or 108 of the 1976 United States
Copyright Act, without either the prior written permission of the Publisher, or authorization through payment of the appropriate per-
copy fee to the Copyright Clearance Center, Inc., 222 Rosewood Drive, Danvers, MA 01923 (website: www.copyright.com). Requests to
the Publisher for permission should be addressed to the Permissions Department, John Wiley & Sons, Inc., 111 River Street, Hoboken,
NJ 07030-5774, (201) 748-6011, fax (201) 748-6008, or online at: www.wiley.com/go/permissions.
Evaluation copies are provided to qualified academics and professionals for review purposes only, for use in their courses during the
next academic year. These copies are licensed and may not be sold or transferred to a third party. Upon completion of the review
period, please return the evaluation copy to Wiley. Return instructions and a free-of-charge return shipping label are available at:
www.wiley.com/go/returnlabel. If you have chosen to adopt this textbook for use in your course, please accept this book as your com-
plimentary desk copy. Outside of the United States, please contact your local representative.
ePUB ISBN 978-1-119-32064-7
The inside back cover will contain printing identification and country of origin if omitted from this page. In addition, if the ISBN on the
back cover differs from the ISBN on this page, the one on the back cover is correct.
Printed in the United States of America.
10 9 8 7 6 5 4 3 2 1
