STUDENT C O M P A N I O N
F U N D A M
b
E N T A L S O F
Biochemistry
LIFE AT b
THE
b MOLECULAR
LEVEL
b
Voet Voet Pratt
F O U R T H E D I T I O N
Akif Uzman Jerry Johnson Joseph Eichberg William Widger
b b b b
Donald Voet Judith G. Voet Charlotte W. Pratt
b b b b b b
,This bpage bis bintentionally bleft bblank
,STUDENT COMPANION TO ACCOMPANY b b b
FUNDAMENTALS OF b
BIOCHEMISTRY
LIFE AT THE MOLECULAR LEVEL
b b b b
Fourth Edition b
Akif Uzman
b
University of Houston b b
Jerry Johnson
b
University of Houston b b
Joseph Eichberg b
University of Houston b b
William Widger b
University of Houston b b
Donald Voet b
University of Pennsylvania b b
Judith G. Voet b b
Swarthmore College b
Charlotte W. Pratt b b
Seattle, Washington b
JOHN bWILEY b& bSONS, bINC
, Cover bDesigner bMadelyn bLesure
bCover bIllustration bNorm
bChristiansenb Cover bPhotos
(Vitruvian bMan): b© bOdysseus/Alamy band b© bDennis bHallinan/Alamy
Molecular bstructures bclockwise bfrom btop: bbased bon bX-ray bstructures bthat bwere brespectively
bdetermined bby bRichard bDickerson band bHorace bDrew, bCaltech; bGerard bBunick, bUniversity bof
bTennessee; bThomas bSteitz, bYale bUniversity; bAlfonso bMondragón, bNorthwestern bUniversity; bVenki
bRamakrishnan, bMRC bLaboratory bof bMolecular bBiology, bCambridge, bU.K.; bAndrew bLeslie band bJohn
bWalker, bMRC bLaboratoryb of bMolecular bBiology, bCambridge, bU.K.
Founded bin b1807, bJohn bWiley b& bSons, bInc. bhas bbeen ba bvalued bsource bof bknowledge band bunderstanding
bfor bmore bthan b200 byears, bhelping bpeople baround bthe bworld bmeet btheir bneeds band bfulfill btheir
baspirations. bOur bcompany bis bbuilt bon ba bfoundation bof bprinciples bthat binclude bresponsibility bto bthe
bcommunities bwe bserve bandb where bwe blive band bwork. bIn b2008, bwe blaunched ba bCorporate bCitizenship
bInitiative, ba bglobal beffort bto baddress bthe benvironmental, bsocial, beconomic, band bethical bchallenges bwe
bface bin bour bbusiness. bAmong bthe bissues bwe bare baddressing bare bcarbon bimpact, bpaper bspecifications band
bprocurement, bethical bconduct bwithin bour bbusiness band bamong bour bvendors, band bcommunity band
bcharitable bsupport. bFor bmore binformation, bplease bvisit bour bwebsite: bwww.wiley.com/go/citizenship.
Copyright b© b2012 John bWiley b& bSons, bInc.
All brights breserved. bNo bpart bof bthis bpublication bmay bbe breproduced, bstored bin ba bretrieval bsystem bor
btransmitted bin bany bform bor bby bany bmeans, belectronic, bmechanical, bphotocopying, brecording, bscanning
bor botherwise, bexcept bas bpermitted bunder bSections b107 bor b108 bof bthe b1976 bUnited bStates bCopyright
bAct, bwithout beither bthe bprior bwritten bpermission bof bthe bPublisher, bor bauthorization bthrough bpayment bof
bthe bappropriate bper-copy bfee bto bthe bCopyright bClearance bCenter, bInc., b222 bRosewood bDrive, bDanvers,
bMA b01923, bwebsite bwww.copyright.com. bRequests bto bthe bPublisher bfor bpermission bshould bbe
baddressed bto btheb
Permissions bDepartment, bJohn bWiley b& bSons, bInc. b111 bRiver bStreet, bHoboken, bNJ
b07030-5774, b(201) b748-6011, bfax b(201) b748-6008, bwebsite bhttp://www.wiley.com/go/permissions.
To border bbooks bor bfor bcustomer bservice, bplease bcall b1-800-CALL bWILEY b(225-5945).
Student bCompanion b(paperback) ISBN-13 b 978-1-1182-1827-3
Printed bin bthe bUnited bStates bof
b Americab10 b 9 b8 b7 b 6 b5 b4 b 3 b2 b1
F U N D A M
b
E N T A L S O F
Biochemistry
LIFE AT b
THE
b MOLECULAR
LEVEL
b
Voet Voet Pratt
F O U R T H E D I T I O N
Akif Uzman Jerry Johnson Joseph Eichberg William Widger
b b b b
Donald Voet Judith G. Voet Charlotte W. Pratt
b b b b b b
,This bpage bis bintentionally bleft bblank
,STUDENT COMPANION TO ACCOMPANY b b b
FUNDAMENTALS OF b
BIOCHEMISTRY
LIFE AT THE MOLECULAR LEVEL
b b b b
Fourth Edition b
Akif Uzman
b
University of Houston b b
Jerry Johnson
b
University of Houston b b
Joseph Eichberg b
University of Houston b b
William Widger b
University of Houston b b
Donald Voet b
University of Pennsylvania b b
Judith G. Voet b b
Swarthmore College b
Charlotte W. Pratt b b
Seattle, Washington b
JOHN bWILEY b& bSONS, bINC
, Cover bDesigner bMadelyn bLesure
bCover bIllustration bNorm
bChristiansenb Cover bPhotos
(Vitruvian bMan): b© bOdysseus/Alamy band b© bDennis bHallinan/Alamy
Molecular bstructures bclockwise bfrom btop: bbased bon bX-ray bstructures bthat bwere brespectively
bdetermined bby bRichard bDickerson band bHorace bDrew, bCaltech; bGerard bBunick, bUniversity bof
bTennessee; bThomas bSteitz, bYale bUniversity; bAlfonso bMondragón, bNorthwestern bUniversity; bVenki
bRamakrishnan, bMRC bLaboratory bof bMolecular bBiology, bCambridge, bU.K.; bAndrew bLeslie band bJohn
bWalker, bMRC bLaboratoryb of bMolecular bBiology, bCambridge, bU.K.
Founded bin b1807, bJohn bWiley b& bSons, bInc. bhas bbeen ba bvalued bsource bof bknowledge band bunderstanding
bfor bmore bthan b200 byears, bhelping bpeople baround bthe bworld bmeet btheir bneeds band bfulfill btheir
baspirations. bOur bcompany bis bbuilt bon ba bfoundation bof bprinciples bthat binclude bresponsibility bto bthe
bcommunities bwe bserve bandb where bwe blive band bwork. bIn b2008, bwe blaunched ba bCorporate bCitizenship
bInitiative, ba bglobal beffort bto baddress bthe benvironmental, bsocial, beconomic, band bethical bchallenges bwe
bface bin bour bbusiness. bAmong bthe bissues bwe bare baddressing bare bcarbon bimpact, bpaper bspecifications band
bprocurement, bethical bconduct bwithin bour bbusiness band bamong bour bvendors, band bcommunity band
bcharitable bsupport. bFor bmore binformation, bplease bvisit bour bwebsite: bwww.wiley.com/go/citizenship.
Copyright b© b2012 John bWiley b& bSons, bInc.
All brights breserved. bNo bpart bof bthis bpublication bmay bbe breproduced, bstored bin ba bretrieval bsystem bor
btransmitted bin bany bform bor bby bany bmeans, belectronic, bmechanical, bphotocopying, brecording, bscanning
bor botherwise, bexcept bas bpermitted bunder bSections b107 bor b108 bof bthe b1976 bUnited bStates bCopyright
bAct, bwithout beither bthe bprior bwritten bpermission bof bthe bPublisher, bor bauthorization bthrough bpayment bof
bthe bappropriate bper-copy bfee bto bthe bCopyright bClearance bCenter, bInc., b222 bRosewood bDrive, bDanvers,
bMA b01923, bwebsite bwww.copyright.com. bRequests bto bthe bPublisher bfor bpermission bshould bbe
baddressed bto btheb
Permissions bDepartment, bJohn bWiley b& bSons, bInc. b111 bRiver bStreet, bHoboken, bNJ
b07030-5774, b(201) b748-6011, bfax b(201) b748-6008, bwebsite bhttp://www.wiley.com/go/permissions.
To border bbooks bor bfor bcustomer bservice, bplease bcall b1-800-CALL bWILEY b(225-5945).
Student bCompanion b(paperback) ISBN-13 b 978-1-1182-1827-3
Printed bin bthe bUnited bStates bof
b Americab10 b 9 b8 b7 b 6 b5 b4 b 3 b2 b1
