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BIOL 318 Evolution – Complete Exam 2 Lecture Notes (Extensive exam-focused content with evolutionary theory, models, and mechanisms; ideal for pre-exam revision) Virginia Commonwealth University, 2025–2026

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This document includes all comprehensive lecture notes for Exam 2 of BIOL 318 at Virginia Commonwealth University, covering Chapters 4–9. Topics span speciation concepts (allopatric, sympatric, parapatric), reproductive isolating mechanisms, genetic incompatibilities, extinction patterns, evolutionary rates and trends, phylogenetic tree construction and interpretation, origins of life (RNA world, protocells, hypercycle model), genome evolution (C-value paradox, G-value paradox), and major evolutionary transitions including symbiosis and multicellularity. A critical resource for mastering midterm content in evolutionary biology.

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Institution
BIOL 318 EVOLUTION
Course
BIOL 318 EVOLUTION

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BIOL 318 EVOLUTION ALL EXAM 2
LECTURE NOTES
(Virginia Commonwealth University)

, Exam 2: Chapter 4, 5 and 10-15

Chapter 4: Speciation- concepts and

mechanisms

Species and Speciation:
• Species are the fundamental unit of biological diversity
• Speciation refers to the processes that produce species
What are species:
• Species have two components
o Spatial
o Temporal
Evolutionary species concept:
• Problems with a “one size” fits all definition of species
• Defines species as:
o Lineage of ancestral-descendant populations which maintains its
identity from other such lineages and which has its own evolutionary
tendencies and historical fate
Morphological species concept:
• Traits evolve through time, visible traits and their relationships to
another=organisms morphology (phenotype)
• Morphologies can vary across separate evolutionary lineages because of
evolutionary processes
• So, species should be able to be defined by differences in their morphologies
• Quantifying morphological diversity is known as phonetics
Biological species concept:
• Evolutionary history affects the accumulation of differences that allow
different morphological types to produce viable offspring
• We can use the ability of two types (populations, different phenotypes,
lineages) to produce viable offspring as a measure of whether or not they are
from the same or different species
• Requires that enough divergence have occurred such that they cannot mate
successfully
o Gene flow among populations decreases diversity and divergence
among populations
• Can use by studying:
o Patterns of gene flow
o Tests of crossing individuals from different populations lacking
connections via gene flow
Phylogenetic Species Concept:
• Evolutionary history results in an underlying tree like structure describing
this history
• Used to help define species:
o Clades are monophyletic groups and monophyletic groups can be
defined by special characters known as synapomorphies

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Institution
BIOL 318 EVOLUTION
Course
BIOL 318 EVOLUTION

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