STUDENT C O M P A N I O N
F U N D A M
,
E N T A L S O F
Biochemistry
LIFE AT ,
THE MOLECULAR
, ,
LEVEL
Voet Voet Pratt
F O U R T H E D I T I O N
Akif Uzman
, Jerry Johnson Joseph Eichberg
, ,
William WidgerDonald Voet
, , ,
Judith G. Voet
, Charlotte W. Pratt
, , ,
,This,page,is,intentionally,left,blank
,STUDENT COMPANION TO ACCOMPANY , , ,
FUNDAMENTALS OF ,
BIOCHEMISTRY
LIFE AT THE MOLECULAR LEVEL
, , , ,
Fourth Edition ,
Akif Uzman
,
University of Houston , ,
Jerry Johnson
,
University of Houston , ,
Joseph Eichberg ,
University of Houston , ,
William Widger ,
University of Houston , ,
Donald Voet ,
University of Pennsylvania , ,
Judith G. Voet , ,
Swarthmore College ,
Charlotte W. Pratt , ,
Seattle, Washington ,
JOHN,WILEY,&,SONS,,INC
, Cover,Designer,Madelyn,Lesure,Cov
er,Illustration,Norm,Christiansen,Cov
er,Photos
(Vitruvian,Man):,©,Odysseus/Alamy,and,©,Dennis,Hallinan/Alamy
Molecular,structures,clockwise,from,top:,based,on,X-
ray,structures,that,were,respectively,determined,by,Richard,Dickerson,and,Horace,Drew,,Caltech;,Gerard,B
unick,,University,of,Tennessee;,Thomas,Steitz,,Yale,University;,Alfonso,Mondragón,,Northwestern,Unive
rsity;,Venki,Ramakrishnan,,MRC,Laboratory,of,Molecular,Biology,,Cambridge,,U.K.;,Andrew,Leslie,and,
John,Walker,,MRC,Laboratory,of,Molecular,Biology,,Cambridge,,U.K.
Founded,in,1807,,John,Wiley,&,Sons,,Inc.,has,been,a,valued,source,of,knowledge,and,understanding,for,more,
than,200,years,,helping,people,around,the,world,meet,their,needs,and,fulfill,their,aspirations.,Our,company,is,
built,on,a,foundation,of,principles,that,include,responsibility,to,the,communities,we,serve,and,where,we,live,a
nd,work.,In,2008,,we,launched,a,Corporate,Citizenship,Initiative,,a,global,effort,to,address,the,environmental,
,social,,economic,,and,ethical,challenges,we,face,in,our,business.,Among,the,issues,we,are,addressing,are,carb
on,impact,,paper,specifications,and,procurement,,ethical,conduct,within,our,business,and,among,our,vendors,
,and,community, and,charitable, support.,For,more,information,,please,visit,our,website:,www.wiley.com/go/c
itizenship.
Copyright,©,2012 John,Wiley,&,Sons,,Inc.
All,rights,reserved.,No,part,of,this,publication,may,be,reproduced,,stored,in,a,retrieval,system,or,transmitted,i
n,any,form,or,by,any,means,,electronic,,mechanical,,photocopying,,recording,,scanning,or,otherwise,,except,
as,permitted,under,Sections,107,or,108,of,the,1976,United,States,Copyright,Act,,without,either,the,prior,writt
en,permission,of,the,Publisher,,or,authorization,through,payment,of,the,appropriate,per-
copy,fee,to,the,Copyright,Clearance,Center,,Inc.,,222,Rosewood,Drive,,Danvers,,MA,01923,,website,www.
copyright.com.,Requests,to,the,Publisher,for,permission,should,be,addressed,to,the,Permissions,Department,
,John,Wiley,&,Sons,,Inc.,111,River,Street,,Hoboken,,NJ,07030-5774,,(201),748-6011,,fax,(201),748-
6008,,website,http://www.wiley.com/go/permissions.
To,order,books,or,for,customer,service,,please,call,1-800-CALL,WILEY,(225-5945).
Student,Companion,(paperback) ISBN-13, 978-1-1182-1827-3
Printed,in,the,United,States,of,America,1
0, 9,8,7, 6,5,4, 3,2,1
F U N D A M
,
E N T A L S O F
Biochemistry
LIFE AT ,
THE MOLECULAR
, ,
LEVEL
Voet Voet Pratt
F O U R T H E D I T I O N
Akif Uzman
, Jerry Johnson Joseph Eichberg
, ,
William WidgerDonald Voet
, , ,
Judith G. Voet
, Charlotte W. Pratt
, , ,
,This,page,is,intentionally,left,blank
,STUDENT COMPANION TO ACCOMPANY , , ,
FUNDAMENTALS OF ,
BIOCHEMISTRY
LIFE AT THE MOLECULAR LEVEL
, , , ,
Fourth Edition ,
Akif Uzman
,
University of Houston , ,
Jerry Johnson
,
University of Houston , ,
Joseph Eichberg ,
University of Houston , ,
William Widger ,
University of Houston , ,
Donald Voet ,
University of Pennsylvania , ,
Judith G. Voet , ,
Swarthmore College ,
Charlotte W. Pratt , ,
Seattle, Washington ,
JOHN,WILEY,&,SONS,,INC
, Cover,Designer,Madelyn,Lesure,Cov
er,Illustration,Norm,Christiansen,Cov
er,Photos
(Vitruvian,Man):,©,Odysseus/Alamy,and,©,Dennis,Hallinan/Alamy
Molecular,structures,clockwise,from,top:,based,on,X-
ray,structures,that,were,respectively,determined,by,Richard,Dickerson,and,Horace,Drew,,Caltech;,Gerard,B
unick,,University,of,Tennessee;,Thomas,Steitz,,Yale,University;,Alfonso,Mondragón,,Northwestern,Unive
rsity;,Venki,Ramakrishnan,,MRC,Laboratory,of,Molecular,Biology,,Cambridge,,U.K.;,Andrew,Leslie,and,
John,Walker,,MRC,Laboratory,of,Molecular,Biology,,Cambridge,,U.K.
Founded,in,1807,,John,Wiley,&,Sons,,Inc.,has,been,a,valued,source,of,knowledge,and,understanding,for,more,
than,200,years,,helping,people,around,the,world,meet,their,needs,and,fulfill,their,aspirations.,Our,company,is,
built,on,a,foundation,of,principles,that,include,responsibility,to,the,communities,we,serve,and,where,we,live,a
nd,work.,In,2008,,we,launched,a,Corporate,Citizenship,Initiative,,a,global,effort,to,address,the,environmental,
,social,,economic,,and,ethical,challenges,we,face,in,our,business.,Among,the,issues,we,are,addressing,are,carb
on,impact,,paper,specifications,and,procurement,,ethical,conduct,within,our,business,and,among,our,vendors,
,and,community, and,charitable, support.,For,more,information,,please,visit,our,website:,www.wiley.com/go/c
itizenship.
Copyright,©,2012 John,Wiley,&,Sons,,Inc.
All,rights,reserved.,No,part,of,this,publication,may,be,reproduced,,stored,in,a,retrieval,system,or,transmitted,i
n,any,form,or,by,any,means,,electronic,,mechanical,,photocopying,,recording,,scanning,or,otherwise,,except,
as,permitted,under,Sections,107,or,108,of,the,1976,United,States,Copyright,Act,,without,either,the,prior,writt
en,permission,of,the,Publisher,,or,authorization,through,payment,of,the,appropriate,per-
copy,fee,to,the,Copyright,Clearance,Center,,Inc.,,222,Rosewood,Drive,,Danvers,,MA,01923,,website,www.
copyright.com.,Requests,to,the,Publisher,for,permission,should,be,addressed,to,the,Permissions,Department,
,John,Wiley,&,Sons,,Inc.,111,River,Street,,Hoboken,,NJ,07030-5774,,(201),748-6011,,fax,(201),748-
6008,,website,http://www.wiley.com/go/permissions.
To,order,books,or,for,customer,service,,please,call,1-800-CALL,WILEY,(225-5945).
Student,Companion,(paperback) ISBN-13, 978-1-1182-1827-3
Printed,in,the,United,States,of,America,1
0, 9,8,7, 6,5,4, 3,2,1
