Arthropods include spiders, scorpions, ticks, mites, crustaceans and insects. Live in aquatic and terrestrial
environments. They are herbivorous, carnivorous or omnivorous.
Why have Arthropods achieved such great diversity and abundance?
- They have a versatile exoskeleton
- Segmentation and appendages provide for more efficient locomotion
- Air piped directly to cells
- Highly developed sensory organs
- Complex behaviour patterns
- Use of diverse resources through metamorphosis
Characteristics of Phylum Arthropoda:
Jointed appendages
Live in aquatic and terrestrial habitats
Some can fly
Free-living and parasitic
Bilateral symmetry
Segmented body divided into tagmata
Triploblastic
Reduced coelom, mostly filled with blood (haemocoel)
Cuticular exoskeleton
Complete digestive system
Complex muscular system
Nervous system consists of a dorsal brain and ganglia
Well-developed sensory organs
Sexes usually separate, with paired reproductive organs and ducts
Usually internal fertilisation
Respiration by body surface, gills, trachea or book lungs
Open circulatory system with dorsal contractile heart, arteries and hemocoel
Subphylum Trilobita
Dorsoventrally flattened bottom dwellers and scavengers
Exoskeleton made of chitin, strengthened in some areas with calcium carbonate
3 tagmata: head (cephalon), trunk and pygidium
Cephalon had a pair of antennae, compound eyes, a mouth and 4 pairs of leglike appendages
No true mouthparts
Each body segment except the last bear a pair of biramous appendages
Subphylum Chelicerata:
Includes horseshoe crabs, spiders, ticks, mites, scorpions and sea spiders. Their bodies are composed of 2 tagmata: a
cephalothorax and an abdomen. They have 6 pairs of cephalothoracic appendages that include a pair of chelicerae, a
pair of pedipalps and 4 pairs of walking legs. They have no antennae. Most chelicerates suck liquid food from their
prey.
Class Merostomata:
Subclass Eurypterida:
- Giant water scorpions - 4 pairs of walking legs on their abdomen
- Head has 6 fused segments - 12 segments and a spike like telson
- Compound eyes - Some had anterior appendages modified into
- Chelicerae crushing jaws
- Pedipalps
, Subclass Xiphosurida: Horseshoe crabs
- Live in shallow water
- Unsegmented horseshoe-shaped carapace and a broad abdomen which has a long telson or tailpiece
- Their cephalothorax bears a pair of chelicerae, one pair of pedipalps and 4 pairs of walking legs
- Their abdomen bears 6 pairs of broad, thin appendages that are fused in the median line
- On 5 abdominal appendages, book lungs occur under the gill opercula
- 2 lateral rudimentary eyes and 2 simple eyes on the carapace
- Swims by means of abdominal plates and can walk with its walking legs
- Feeds at night on worms and small molluscs, which it seizes with its chelicerae and walking legs
- During mating season, they come to shore at high tide to mate. A female, burrows into sand and lays eggs. Males
add sperm to the eggs and the female covers it with sand. Eggs are warmed by the sun.
- Larva are segmented
Class Pycnogonida: sea spiders
- Small thin bodies
- 4 pairs of long thin walking legs
- Segments are duplicated in some, so they possess 5 or 6 pairs of legs
- Some males have a pair of legs (ovigers) on which they carry developing eggs
- Equipped with chelicerae and pedipalps
- The small head (cephalon) has a raised projection with 2 pairs of simple eyes
- Mouth at the tip of a long proboscis, which sucks juices from soft-bodied animals
- Circulatory system is limited to a simple dorsal heart
- No excretory or respiratory system
- Long thin body and legs provide large surface area in proportion to body volume for diffusion of gas and wastes
- Due to the small size of the body, the digestive system and gonads have branches that extend into the legs
Class Arachnida:
- Common in warm dry areas
- 2 tagmata: a cephalothorax and an abdomen, which may or may not be segmented
- The abdomen houses reproductive and respiratory organs
- Cephalothorax bears a pair of chelicerae, pair of pedipalps and 4 pairs of walking legs
- Mostly predaceous and have chelicerae modified into fangs and pedipalps modified into claws
- Strong sucking pharynx
- Spinning glands
Order Araneae: Spiders
The spider body is compact
Consists of a cephalothorax and abdomen, both unsegmented and joined by a slender pedicel
Pair of chelicerae which have terminal fangs through which ducts run from venom glands
Pair of leg-like pedipalps which have sensory function and are used by males to transfer sperm
4 pairs of walking legs which terminate in claws
All spiders are predaceous. They seize prey with their chelicerae, inject venom, liquify the prey’s tissues
with digestive fluid and sucks the resulting broth into its stomach
Respire using book lungs or a system of internal tubes called tracheae.
Book lungs open to the outside through a slit in the exoskeleton. Internal to the slit are many thin-walled
chambers filled with blood. Between a pair of chambers is a cavity where air enters
Excretory system of Malpighian tubes and specialised resorptive cells in the intestinal epithelium
8 simple eyes
Web-spinning habits:
2-3 pairs of spinnerets containing hundreds of microscopic tubes run to abdominal silk glands
A scleroprotein secretion emitted as a liquid from the spinnerets hardens to form a silk thread
Threads of spider silk are stronger than steel threads of the same diameter
Kind of net spun varies among species
Use nets for nest lining, sperm webs, egg sacs, bridge lines, warning threads, molting threads,
attachment discs, nursery webs and trapping of prey